Iranian Supreme Leader Ayatollah Ali Khamenei has warned Israel and the United States of “a crushing response” for actions against Iran and its allies, according to state media.

Khamenei, 85, made the remarks on Saturday while addressing students ahead of the anniversary of the 1979 takeover of the US embassy in Tehran by hardline students – which cemented the decades-long enmity between Tehran and Washington that persists today.

“The enemies, whether the Zionist regime or the United States of America, will definitely receive a crushing response to what they are doing to Iran and the Iranian nation and to the resistance front,” Khamenei said in the capital, Tehran, also referring to Iran-aligned armed groups that include Yemen’s Houthis, Lebanon’s Hezbollah and the Palestinian Hamas.

The supreme leader did not elaborate on the timing of any attack, or the scope.

He had previously struck a more cautious approach, saying officials would weigh Iran’s response and that Israel’s attack “should not be exaggerated nor downplayed”, after the Israeli military launched strikes last week on military bases in Iran, hitting about 20 sites over several hours in Ilam, Khuzestan and Tehran.

Khamenei on Saturday met with university students to mark Students’ Day, which commemorates a November 4, 1978, incident in which Iranian soldiers opened fire on students protesting the rule of the shah at Tehran University.

The US military operates throughout the Middle East, with some troops now manning a Terminal High Altitude Area Defense, or THAAD, battery in Israel.

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